Protection Against Osmotic Stress by cGMP-Mediated Myosin Phosphorylation

DISRUPTION Threonine Cytoplasm HEAVY-CHAIN PHOSPHORYLATION CYCLIC-AMP SIGNAL-TRANSDUCTION Myosins 03 medical and health sciences Osmotic Pressure Animals Dictyostelium Pseudopodia Phosphorylation Cyclic GMP 0303 health sciences DICTYOSTELIUM-DISCOIDEUM Water-Electrolyte Balance GENE Actins REPLACEMENT Actin Cytoskeleton Glucose MUTANTS Guanylate Cyclase LACKING ATPASE ACTIVITY
DOI: 10.1126/science.271.5246.207 Publication Date: 2006-10-27T18:30:41Z
ABSTRACT
Conventional myosin functions universally as a generator of motive force in eukaryotic cells. Analysis of mutants of the microorganism Dictyostelium discoideum revealed that myosin also provides resistance against high external osmolarities. An osmo-induced increase of intracellular guanosine 3′,5′-monophosphate was shown to mediate phosphorylation of three threonine residues on the myosin tail, which caused a relocalization of myosin required to resist osmotic stress. This redistribution of myosin allowed cells to adopt a spherical shape and may provide physical strength to withstand extensive cell shrinkage in high osmolarities.
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